Published in Agron. J. 96:1651-1659 (2004).
© American Society of Agronomy
677 S. Segoe Rd., Madison, WI 53711 USA
Tillage and Cropping Systems
Soil Structural Disturbance Effects on Crop Yields and Soil Properties in a No-Till Production System
M. Díaz-Zoritaa,*,
J. H. Groveb,
L. Murdockb,
J. Herbeckb and
E. Perfectc
a Dep. of Plant Prod., Faculty of Agron., Univ. of Buenos Aires, 1417, Av. San Martín 4457, Buenos Aires, Argentina, and Nitragin Argentina S.A., Calle 10 y 11, Parque Industrial Pilar, 1629, Pilar, Buenos Aires, Argentina
b Dep. of Agron., Univ. of Kentucky, Lexington, KY 40546-0091
c Dep. of Earth and Planetary Sci., Univ. of Tennessee, Knoxville, TN 37996-1410
* Corresponding author (mdzorita{at}speedy.com.ar)
Received for publication December 14, 2003.
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ABSTRACT
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The development of well-structured soils is a goal for achieving sustainable and productive agricultural systems. Nevertheless, the maintenance of soil structure in continuous no-till (NT) soils has sometimes been thought to induce soil conditions that are detrimental to crop yields. The objectives of this research were to characterize the effects of periodic tillage disruption in otherwise NT systems on soil properties and the yields of winter wheat (Triticum aestivum L.), double-cropped soybean [Glycine max (L.) Merr.], and maize (Zea mays L.) in rotation and to determine if soil structural changes occurring in tilled soils are independent of changes in other soil properties. A field experiment was established in 1992 on a Huntington silt loam soil (Fluventic Hapludoll) at the University of Kentucky Research and Education Center in Princeton (KY) under a NT crop sequence with two seedbed preparation methods for winter wheat, (a) NT or (b) chisel plus disk tillage (Till). In fall 2000, similar soil chemical properties were observed between disrupted and continuous NT systems over the 0- to 20-cm layer. The geometric mean diameter of dry fragments and the soil water content retained between 0.0003 and 0.03 MPa water potential was greater in NT soils than in soils tilled for winter wheat. Tillage for winter wheat enhanced winter wheat yields (4.2% increase), mostly under low-yielding conditions, but it resulted in a reduction of subsequent summer crop yields (i.e., 3.7% for soybean and 7.0% for maize). The yields obtained in our study translate to an economic benefit for the continuous NT system. Net returns per hectare were estimated to be $73 higher for the winter wheat/double-crop soybeanmaize rotation under NT than under Till treatments. The differences in maize yields between NT and tilled treatments were attributed to a better water supply in NT soil due to the maintenance of a larger number of mesopores and a great hydraulic conductivity. In the absence of significant changes in other physicochemical properties, periodic tillage appears to disrupt soil structure, which negatively affects crop productivity.
Abbreviations: GMD, geometric mean diameter LogGSD, log of the geometric standard deviation NT, no-till SWC, soil water content Till, chisel plus disk tillage TN, total nitrogen TOC, total organic carbon
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INTRODUCTION
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THE SPATIAL ARRANGEMENT of voids and solids is involved in many of the processes occurring in agricultural soils that influence crop productivity (Kay and Angers, 1999). Aeration, imbibition of water, soil temperature change, and seedsoil contact are controlled by the soil structure of seedbeds and affect germination and establishment of crops (Hewitt and Dexter, 1979; Braunack and Dexter, 1989a, 1989b; Nasr and Selles, 1995). Root development is favored by the presence of continuous pores allowing soil exploitation by the crop (Russell, 1977).
The development of structure in soils is also important for controlling wind and/or water erosion processes (Lal and Elliott, 1994; Skidmore et al., 1994). For example, the presence of large stable soil fragments in seedbeds promotes the occurrence of surface roughness, reducing soil erodibility by wind (Chepil, 1953) as well as water erosion (Deizman et al., 1987). Well-structured soils have higher infiltration rates and can maintain their pore system in a stable state longer than soils formed by weak structural units (Kay, 1990).
In agricultural soils, tillage and traffic operations are major factors involved in soil structure degradation through fragmentation and compaction processes (Kay, 1990). Fragmentation involves the break down of large structural units into smaller fragments. Compaction involves a reduction in the void space. In both processes, the final soil structural status will depend on the amount of mechanical stress applied and soil properties [e.g., organic matter, texture, soil water content (SWC)].
The surficial accumulation of soil organic materials favored by minimum tillage disruption in conservation tillage systems is recognized to be a major factor contributing to the development of structural units similar to those observed under sod or native conditions (Lal et al., 1994). No-tillage practices have been reported to maintain and sometimes enhance soil aggregation when compared with other tillage practices (Mahboubi and Lal, 1998; Díaz-Zorita, 1999). Another advantage of NT systems is conservation and storage of soil water due to reduced water evaporation and increased infiltration in the presence of surface mulches (Mahboubi et al., 1993; Unger and Jones, 1994).
The maintenance of continuous NT systems may produce soil conditions (higher soil compaction, lower soil temperatures) that reduce crop productivity (Lal et al., 1994). For example, greater soil penetration resistance and bulk density levels are commonly observed in NT soils compared with frequently tilled soils (Van Ouwerkerk and Boone, 1970; Kruger, 1996). Several researchers have observed winter wheat grain yield reductions when planted after maize crops in NT systems and attributed this negative effect to either prolonging low temperatures during spring or to poor stand establishment in the presence of large amounts of residues (e.g., Mead and Chan, 1988; Kiger and Grove, 1999). Winter wheat plant, tiller, and spike densities increased when the distance between maize residue and the wheat row was increased (Kiger and Grove, 1999). Thus, tilled winter wheat seedbeds are the common practice currently used by producers to avoid winter wheat grain yield reductions.
Pierce et al. (1994) studied the effects of periodic moldboard plowing on soil properties of soils in NT farming systems in eastern Minnesota. They observed that plowing, compared with NT, decreased bulk density and increased total and macroporosity levels. Within 4 to 5 yr after plowing, the effects of plowing were dissipated, and the soil was physically similar to NT. Tilling a soil previously subjected to NT redistributed soil nutrients within the plow layer, but the surface stratification of NT was not entirely lost. Residual effects of periodic plowing were minimal for soil physical properties but significant for chemical properties including organic C levels (Pierce and Fortin, 1997).
The information available supporting the regular implementation of tillage practices in otherwise NT systems is not abundant (Pidgeon and Soane, 1977; Campbell et al., 1984). Varsa et al. (1997) and Díaz-Zorita (2000) showed that periodic loosening of NT soils with deep-tillage operations does not always result in increased crop yields. West et al. (1996) concluded that deep loosening improved maize yields on dark, poorly drained soils but not on well-drained soils where NT is traditionally well adapted.
The objectives of this research were (i) to characterize the effects of periodic tillage disruption in otherwise NT systems on soil properties and crop productivity in a winter wheatdouble-crop soybeanmaize rotation in western Kentucky and (ii) to determine if soil structural changes occurring in tilled soils are independent of changes in other soil properties.
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MATERIALS AND METHODS
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Site Description and Experiment Design
The experiment was established in the fall of 1992 at the University of Kentucky Research and Education Center in Princeton (KY) on a Huntington silt loam soil (Fluventic Hapludoll) that is moderately well drained. The study utilized a common crop rotation in Kentucky: maizewinter wheatdouble-crop soybean. The double-crop soybean was planted after winter wheat in the same production year. The experiment was established on a site where NT agricultural practices had been used for more than 20 yr.
Two winter wheat seedbed planting systems, NT winter wheat and tilled winter wheat (Till), were compared using a complete randomized block design in plots 9 m long by 3 m wide with three replicates. No-till winter wheat was planted directly into mechanically shredded maize stalks with a NT drill. Tilled winter wheat plots were chisel-plowed and disked twice before planting. Maize and double-crop soybean crops were always planted with NT practices in both winter wheat planting systems (NT and Till) so that the only tillage difference in the entire rotation was tillage for winter wheat. Each year, the three crops were planted at an optimum time: winter wheat, early to mid-October; double-crop soybean, mid-June; and maize, early April. Crop cultivars changed from year to year. Lime and N, P, and K fertilizer applications were made according to recommendations (Anonymous, 2003) and were similar for both treatments (NT and Till).
Crop Measurements and Weather Information
Maize, soybean, and winter wheat grain yields were determined yearly since 1992 until 2000 by harvesting the center rows of each plot with a plot combine. All grain yields were adjusted to a moisture content of 130 g kg1. Maize grain yields for the 1996 season were not reported because of failures during the harvest operation.
Daily precipitation records for the studied period, from the weather station located at the University of Kentucky Research and Education Center in Princeton (KY), were downloaded from the University of Kentucky Weather Center (http://wwwagwx.ca.uky.edu; verified 22 July 2004).
Soil Measurements
Composite soil samples from the midinterrow space of each plot, where low machinery traffic occurs, were taken in the spring of 2000, 20 and 8 mo after the fourth tillage operation (spring 1998 and 1999). At soil sampling, maize and soybean crops were within the first month after planting.
Total Organic Carbon, Nitrogen, pH, and Soil Test Phosphorus and Potassium
Twelve composite soil samples from the 0- to 10- and 10- to 20-cm layers were taken from each plot using a 1.6-cm-diam. cylindrical core sampler. Soil total nitrogen (TN) and total organic carbon (TOC) concentrations were determined by dry combustion using a LECO CN-2000 (Leco Corp., St. Joseph, MI) analyzer (Tabatabai and Bremner, 1970). Soil water pH in a 1:1 slurry was measured with deionized water electrometrically (Thomas, 1996). Soil test extractable P and K were determined according to the Mehlich-III procedure (Mehlich, 1984).
Bulk Density and Saturated Hydraulic Conductivity
Duplicate soil core samples were taken in the 0- to 10- and 10- to 20-cm soil layers with a 5.2-cm-diam. and 137.4-cm3 cylinder for measuring soil oven dry bulk density (Blake and Hartge, 1986). The mean coefficient of variation between the duplicates was 16.3%. Only in the plots cultivated with maize, 20 mo after the last tillage operation, a separate triplicate set of samples was taken from the 0- to 10-cm soil layer for determining saturated hydraulic conductivity using a constant head permeameter (Klute and Dirksen, 1986). The average coefficient of variation between triplicates was 27.8%.
Dry Fragment Size Distribution
Duplicate soil samples in the 0- to 10- and the 10- to 20-cm layers were taken with a 5.2-cm-diam. cylinder when the gravimetric SWC averaged 219 g kg1. No significant differences in SWC between tillage treatments, 212 g kg1 for NT and 226 g kg1 for Till, were observed. Each field-moist sample was dropped from a height of 1.6 m over an aluminum tray to induce fragmentation and air-dried for at least 14 d before sieving. The mean SWC after air drying was 14.1 g kg1. Dry sieving was performed with a Fritsch vertical vibratory sieve shaker (model Analysett-3, Fritsch, Idar-Oberstein, Germany) for 30 s using an oscillation amplitude of 2 mm and a frequency of approximately 50 Hz (Díaz-Zorita et al., 2000). The dry fragment size distribution was determined using a nest of sieves that consisted of 13 aperture sizes between 16.00 and 0.85 mm using the mass of fragments remaining on each sieve after sieving to calculate the distribution of fragments normalized with respect to the total mass. The fragment size distributions were characterized using the geometric mean diameter (GMD) and the log of the geometric standard deviation (LogGSD) parameters from the cumulative log-normal distribution equation fitted using the NLIN procedure of PC-SAS (SAS Inst., 1997). The upper size of fragments was estimated as the maximum length of the sampled soil (i.e., 100 mm).
Soil Water Retention Curves
Composite-disturbed samples from the 0- to 10-cm layer of the soil were taken with a 1.6-cm-diam. cylinder and used for constructing soil water retention curves. Dew point water activity meters (WP4 Dewpoint PotentiaMeter and AquaLab, Decagon Devices, Inc., Pullman, WA) were used to measure soil water tension (
) over a range of SWCs from approximately 100 kPa to oven dryness (Gee et al., 1992). The corresponding gravimetric water contents were measured, and the data was fitted to the Ross et al. (1991) soil water retention model using Eq. [1],
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where
d is the tension at oven dryness, c is an empirical constant related to the pore-size distribution, and A (Eq. [2]) is a compound constant related to the SWC at saturation (SWCsat), tension at the air entry value (
a),
d, and c,
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The A,
d, and c in Eq. [1] were estimated using the NLIN procedure of PC-SAS (SAS Inst., 1997). All of the fits converged and had adjusted r2 values greater than 0.98.
Soil water retention curves were also measured using disturbed and undisturbed samples. A knife was used to cut a 0.5-cm-thick rectangular soil block from a depth between 2 and 3 cm below the surface. This was performed 1 d after a rainfall, permitting excavating and trimming to a cross-sectional area of 1 cm2 with minimal disturbance. For transportation, the undisturbed samples were placed into the same plastic containers provided for making the water tension measurements with the dew point water activity meter. Additional samples, taken from directly adjacent soil, were placed in a plastic bag and broken by hand into small fragments under field moisture conditions. Equation [1] was fitted to the two data sets using the NLIN procedure of PC-SAS (SAS Inst., 1997), yielding estimates of A,
d, and c. All of the fits converged and had adjusted r2 values greater than 0.97.
Soil Water Content
In 1999 and 2000 maize growing season, beginning one month after planting, 15 composite samples were taken every 2 wk from the 0- to 15-, 15- to 30-, and 30- to 45-cm layers of the soil with a 1.6-cm-diam. cylinder from all plots planted with maize and used to determine the gravimetric SWC.
Penetration Resistance
In each plot in its second year after tillage, 4 d after a rainfall, six sampling points were located along the midinterrow for measuring the soil strength in the 0- to 30-cm layer at 1-cm intervals using a recording cone penetrometer (Eijelkamp Agrisearch Equipment, Giesbeek, the Netherlands). The gravimetric SWC, for the 0- to 10- and 10- to 20-cm soil layers, was determined by compositing samples taken close to each point of penetration.
Data Analysis
Analysis of variance was performed for interpreting soil management effects on seasonal crop grain yield for a randomized block design using the GLM procedure and protected LSD test of PC-SAS (SAS Inst., 1997) at a 95% level of confidence. Seasonal mean yield effects on treatment (NT and Till) yields were evaluated comparing regression lines (Snedecor and Cochran, 1989) between both variables for each crop (Analytical Software, 2000).
For each crop sequence (winter wheatsoybean or maize plots), soil management (NT and Till) effects on TN, TOC, pH, soil test P and K, bulk density GMD and LogGSD parameters, and penetration resistance levels were analyzed using analysis of variance (SAS Inst., 1997) for a randomized block design split by depth of sampling. Saturated hydraulic conductivity and soil water retention parameters were evaluated using one-way analysis of the variance (SAS Inst., 1997).
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RESULTS AND DISCUSSION
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Tillage Effects on Crop Yields
Mean annual winter wheat grain yield ranged from 3624 to 6858 kg ha1, and in three out of the eight seasons, tillage for winter wheat enhanced grain yields with respect to continuous NT management (Table 1). Averaged over the eight seasons, winter wheat grain yields in NT were 4.2% lower than in the treatments with tillage. Excessive surface compaction in 19921993 and below-normal March temperatures in the 19951996 and 19971998 seasons explained most of these differences in yield (Murdock et al., 2000). The difference in winter wheat grain yield between Till and NT treatments increased when the average March temperature decreased (Fig. 1).
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Table 1. Yearly effect of two soil management practices on winter wheat, double-cropped soybean, and maize yields. Different letters within a column indicate significant differences (P < 0.05) due to tillage systems for a year.
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Fig. 1. Relationship between average March temperature and winter wheat grain yield difference between no-till and tilled wheat seedbed conditions.
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When plotted as a function of the average yields for each season, annual winter wheat production depended on tillage management. Linear models fitted between treatment and seasonal yields showed statistically different slopes (P < 0.0001), indicating that the difference in productivity between Till and NT management varies depending on the seasonal productivity (Fig. 2a). The lower the average seasonal yield, the greater the positive effect of tillage on winter wheat yield. No statistical relationships between the number of seasons under NT practices and the difference, or the ratio, in winter wheat grain yields between Till and NT were observed.

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Fig. 2. Seasonal productivity effect on crop grain yields due to soil tillage for winter wheat (Till) or continuous no-tillage (NT) in a winter wheatdouble-cropped soybeanmaize rotation in Princeton (KY).
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Mean double-cropped soybean yields ranged from 892 to 3267 kg ha1, with significantly higher yields under continuous NT management in only one out of the seven studied seasons (Table 1). Averaged over the seasons, double-cropped soybean yields were 3.7% higher in continuous NT management than in Till for winter wheat treatments. The slopes of the linear relationships between treatment yields and seasonal yields were not statistically different (P = 0.44). The intercept of the regression line for the NT treatment was significantly greater (P = 0.04) than for the Till (Fig. 2b). These results suggest that tillage for winter wheat induced smaller mean double-crop soybean yields than continuous NT management and that this effect was independent of the productivity level of the season. There were no statistical relationships between the number of seasons under NT practices or the amount of rainfall in July and August and the difference, or the ratio, in double-cropped soybean grain yields between Till and NT.
Mean maize grain yields ranged from 9277 to 11903 kg ha1, and in most of the seasons, four out of six, the yields under continuous NT management were significantly greater than those where tillage was done before winter wheat planting (Table 1). Averaged over all six seasons, maize under NT management yielded 7.0% more than in Till treatments. The slopes of the linear relationship between the average seasonal yield and treatment yields were not statistically different (P = 0.60). However, a statistically greater intercept (P = 0.02) was determined for continuous NT maize compared with Till for winter wheat treatments (Fig. 2c).
Maize grain yields were increased by continuous NT management, independent of seasonal productivity. Nevertheless, greater enhancement of maize yields under continuous NT management occurred in 1994 and 1999 seasons with greater water deficiencies than when adequate rainfall occurred (Table 2). The percentage of yield change under continuous NT with respect to Till management was negatively related to the total amount of rainfall in May [yield change (%) = 22.3 0.13 x rainfall (mm), r2 = 0.76, P = 0.023].
No statistical relationships between the number of seasons under NT practices and the difference, or the ratio, in maize grain yields between Till and NT were observed.
The yields obtained in our study translate to an economic benefit for the continuous NT system. This conclusion was based on the average grain yields, total cost, and grain prices for farmers in western Kentucky for the studied period (1991 to 2000). For the complete maizewinter wheatdouble-crop soybean sequence, the production costs were only different for winter wheat. We assume that the cost of chisel tillage, with one secondary disking, was about $50 ha1, including the fuel and the maintenance of a larger tractor for primary and secondary tillage. The cost of herbicide use, including spraying, was estimated to be $40 ha1. Thus, net returns for the complete maizewinter wheatdouble-crop soybean sequence were estimated to be $73 ha1 higher under NT than under Till treatments. This comparison does not include government payments and promotions.
Tillage Effect on Soil Properties
Soil test pH, P, K, TOC, TN, and bulk density levels, either 8 or 20 months after the tillage operation in 1999, were similar between NT and Till treatments and independent of the sampling depth (Table 3). Mean values of these properties, averaged by management treatment or by sampling depth, are presented in Table 4. The absence of significant differences in pH, P, or K levels was expected because they were treated equally in both tillage treatments. The TOC and TN levels were probably the same for both treatments because the tillage operations for winter wheat were performed using tillage tools without soil inversion and in the fall when low temperatures as well as high SWCs are the rule. Conservation tillage practices, those that maintain at least 30% soil surface coverage with harvest residues at planting, promote surface accumulation of C because of reduced mineralization and erosion (Thomas, 1985; Golchin et al., 1994; Buschiazzo et al., 1998). Furthermore, soil mineralization is reduced with a reduction in soil temperature and/or aeration (Skopp et al., 1990; Paul and Clark, 1996). Bulk density changes in agricultural soils with similar textures were related to machinery traffic and losses of organic matter after tillage operations (Kay, 1990). In this study, the lack of statistically significant differences in bulk density between tillage treatments can be explained by the fact that soil samples were taken in the interrow space where low machinery traffic occurred and that there were no significant changes in organic matter levels.
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Table 3. Results of analyses of variance for soil properties in a Huntington silt loam soil under two tillage management systems (chisel plus disk tillage vs. no-tillage). P = soil test P, K = soil test K, TOC = total organic carbon, TN = total nitrogen, and BD = bulk density.
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Table 4. Mean soil properties at two depths of a Huntington silt loam soil under two tillage management systems. P = soil test P, K = soil test K, TOC = total organic carbon, TN = total nitrogen, and BD = bulk density.
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The soil penetration resistance levels, averaged over the sampled profile, were 35% greater for the NT soils (1.24 MPa) than the Till soils (0.92 MPa). The gravimetric SWCs at sampling, averaged over both depths, were not related with the penetration resistance values and were similar between NT and Till treatments, 273 and 289 g kg1, respectively. No statistically significant interactions between tillage treatments and sampling depth were observed. Significantly greater soil strength/compaction was observed for the 17- to 20-cm layer in NT than Till treatments (Fig. 3). The greater strength observed under NT soils is attributed to the presence of more cohesive soil, root networks, and higher density associated with the reorientation of charged surfaces favored by the absence of tillage disruption (Kay, 1990). In this study, the penetration resistance values were always smaller than reported critical levels for normal root growth and crop productivity (Greacen and Oh, 1972; Ehlers et al., 1983; Boone et al., 1986; Gupta and Allmaras, 1987). Consequently, differences in soil strength between tillage treatments do not directly explain the differences in summer crop yields where maize yields were generally greater on NT soils with the higher penetration resistance values.

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Fig. 3. Effect of two management practices, continuous no-tillage (NT) and tillage (Till) for winter wheat, in a winter wheatdouble-cropped soybeanmaize crop sequence on soil penetration resistance (PR) levels in a Huntington silt loam soil. * and ** indicate significant differences in PR between tillage treatments at the 90 and 95% levels of confidence, respectively.
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Almost 2 yr after performing tillage for winter wheat, tillage resulted in significantly (p = 0.05) smaller saturated hydraulic conductivity in the 0- to 10-cm layer compared with soils under NT practices, 5.1 and 16.4 mm h1, respectively. Similar results have been described in other tillage experiments and the response attributed to the greater macropore continuity in NT systems as well as the stability of the pore system in the presence of moving water (Thomas, 1985; Mahboubi et al., 1993).
No significant interaction between sampling depth and tillage treatments on fragment size distribution was observed. Averaged over the 0- to 20-cm layer, significantly larger GMDs of dry fragments were found under NT (13.2 mm) than under Till (11.1 mm). No significant effect of tillage practices on the LogGSD of dry fragments was observed. The occurrence of large-sized fragments indicates that organic and inorganic bonds between primary soil particles were stronger under NT systems.
Soil water retention curves based on disturbed samples were different for the two tillage treatments (Fig. 4). In this experiment, because of the lack of difference in bulk density between treatments, differences in gravimetric water content can be interpreted mostly in terms of pore size distribution. At 0.01 MPa of water tension, equivalent to pores 30 µm in diameter, the gravimetric SWC was significantly greater in soils under continuous NT (0.265 g g1) than in those under tillage for winter wheat (0.215 g g1). No significant differences (P > 0.10) between the two tillage treatments were observed in the gravimetric SWC at a water tension of 1.5 MPa, equivalent to pores with 0.20-µm diameter. The proportion of mesopores, i.e., pores with equivalent diameter range from 10 to 1000 µm (Luxmoore, 1981), was 23% greater in NT soils (0.189 cm3 cm3) than in soils tilled once every 2 yr for winter wheat (0.154 cm3 cm3).

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Fig. 4. Soil water retention curves for a Huntington silt loam soil under two soil management systems using disturbed soil samples. NT = continuous no-tillage, and Till or T = seedbed tillage for winter wheatnumbers 1 to 3 identify the three field replications. Solid lines indicate predicted relationship from the best fit of Eq. [1], and the model parameters are in Table 5. SWC, soil water content.
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Unger (1975) observed that large errors can be introduced when estimates of field soil water relations are based on water retention measurements using sieved soil in the laboratory. We also observed that sample pretreatment affected the relationship between gravimetric SWC and water tension, mostly for NT soils. In general, greater SWC levels were determined using undisturbed rather than disturbed samples (Table 5). The difference between handling treatments was larger at low water tension levels, in the presence of large pores, than at high water tensions. Elrick and Tanner (1955) evaluated the influence of sample pretreatment on soil moisture retention and reported that 2-mm sieving of medium-textured soils decreased moisture retention compared with core samples when the measurements were performed at tensions greater than 0.1 MPa. Sample pretreatment (disturbed vs. undisturbed) was not important when samples came from the tilled winter wheat treatment. It is hypothesized that the maintenance of NT practices enhances the proportion of small-sized pores resistant to physical disruption during sieving and consequently raises their contribution to soil water storage. In Table 5, estimates for the predicted soil water retention curve fitted using Eq. [1] are reported. Because of the variability in these parameters, further research is needed to verify the impact of soil disturbance on the relationship between gravimetric water content and water potential determined using water activity meters.
Under field conditions, the greater SWC in NT soils has been attributed mostly to reduced evaporation due to the presence of surface residue cover, an "organic mulch" (Blevins et al., 1971; Unger, 1994; Tanaka and Anderson, 1997). In 1999, greater SWCs were found under continuous NT management during most of the maize growing season (Fig. 5). No significant differences were found the following season (Fig. 4) when greater rainfall occurred (Table 2). Soil residue cover 20 months after the last tillage for winter wheat was similar between management treatments (data not shown), implying similar protection against evaporative losses between tillage systems. Thus, these results suggest that differences in soil structure resulted in greater water storage in continuous NT soils and improved drought resistance.

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Fig. 5. Effect of two management systems, continuous no-tillage (NT) and tillage (Till), for wheat on the soil water content (0- to 45-cm layer) of a Huntington silt loam soil planted to maize. * indicates significant differences in soil water content between tillage treatments at the 95% level of confidence.
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CONCLUSIONS
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The disruption of soil structure in NT agricultural soils, without causing significant changes in other soil properties, does affect crop productivity. Seedbed tillage for winter wheat enhanced winter wheat grain yields, mostly under low-yielding conditions, because of decreased compaction and warmer soil temperatures compared with NT winter wheat. However, it induced large negative effects on subsequent summer crop yields. The yields obtained in our study translate to an economic benefit for the continuous NT system. Net returns per hectare were estimated to be $73 higher for the winter wheatdouble-crop soybeanmaize rotation under NT than under Till (only for wheat) treatments.
The difference in maize yields between continuous NT and Till for winter wheat soils was greater in seasons with few rainfall events, indicating the importance of maintaining good soil structure to facilitate an adequate water supply for the crop. Greater water storage was observed under continuous NT mostly because of the contributions of a greater percentage of mesopores and pore continuity (greater Ksat values).
These results suggest that soil structure could play a role in reducing losses of soil water to the atmosphere, probably due to differences in the proportion of mesopores and the continuity of macropores. In continuous NT soils, greater soil water conservation can be attributed to the cohesion of small soil aggregates with smaller intra-aggregate pore sizes that favor water storage, thereby reducing evaporation. In addition, a lack of tillage disruption allows the stabilization of interaggregate pores with a range of sizes that significantly contribute toward greater soil water infiltration and storage. The period without tillage in the tilled winter wheat treatment, 2 yr, was not long enough to permit soil structure recovery. Similar results have been reported elsewhere, indicating that the benefits of NT soils, in terms of soil structurerelated processes, require continuity in NT soil management over much longer periods of time (Voorhees and Lindstrom, 1984; Bergh et al., 2000; Karunatilake et al., 2000; McGarry et al., 2000).
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ACKNOWLEDGMENTS
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The authors appreciate the financial support provided by the Kentucky Small Grain Growers Association. We also thank J.H. James for his assistance with field operations.
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